I don't think their is a zombie queen. But going by what I'm seeing throughout the podcast, the "zombies" appear to have a "Pack" mentality. Think of it as a wolf pack I guess, you have an alpha male, female, whatever. Basically one that is in charge of everything. You have your scouts which look for the prey report back to the pack, and then the rest of them round up and hunt. The alpha coordinates by setting the trap, ambush, (yes wolves do this too) and then you corral the prey into the area and strike. Basically what happened in Chapter 21-1. The alpha is usually either the smartest or the strongest, which in this case "Ink" appears to be both. Whatever this "virus" or whatever it turns out to be is, appears to take the human back to their primal stage. And just like the flow of evolution they are learning and adapting to their surroundings, using both whatever intelligence they once had, or they are being taught by another.